Postgraduate study
Edinburgh: Extraordinary futures await.

Integrative Physiology (Biomedical Sciences - Centre for Discovery Brain Sciences) PhD, MScR

Awards: PhD, MScR

Study modes: Full-time, Part-time

Funding opportunities

Not only has the PhD given me the opportunity to learn a wide range of lab techniques that will be beneficial throughout my career, it has also taught me how to be resilient and has helped me to build on my ability to confidently communicate with fellow researchers.

Natalie Courtney PhD Integrative Physiology 2015-2019
Natalie Courtney               PhD Integrative Physiology 2015-2019

The Centre for Discovery Brain Sciences (CDBS) carries out research at molecular, cellular, systems and behavioural levels to understand fundamental mechanisms and pathways relevant to brain and body function in health and disease.

CDBS investigators exploit rapid advances in the enabling technologies available from genomics, proteomics, imaging, informatics, and in-vivo analysis to understand the function of gene products at the cell, organ and whole-animal level. They exploit the most appropriate model organisms/systems to investigate the delicate balance between high biomedical relevance (for example human, mouse, rat) and high genetic power (such as C. elegans, drosophila or zebrafish).

Research focuses on understanding fundamental mechanisms and pathways relevant to human function in health and disease across the life course.

PhD degree

We normally have about 60 Integrative Physiology PhD students on programme at any given time.

Postgraduate students carry out their studies within a research group under the supervision of an academic staff member (their ‘primary’ supervisor). In addition, students are assigned a personal PhD thesis committee comprising their primary supervisor, one or more assistant supervisors and another academic staff member, who provide advice and mentoring, and monitor progress. Supervisors maintain the highest standards of research training with a strong research output in leading international journals.

In their first year, students have the opportunity of participating in relevant taught modules of a number of associated MSc courses. Students attend research seminars and can attend generic skills training programmes provided by the Institute of Academic Development. Postgraduates also have the opportunity to act as demonstrators for undergraduate teaching.

All students are encouraged to present their findings at national and international conferences and to publish their findings in international journals, in advance of submission of their PhD theses for examination.

MSc by Research

Our MSc by Research in Integrative Physiology is a full-time 1-year research project done under the supervision of a CDBS Researcher. This programme has no formally taught component and is therefore only suitable for highly motivated students with a clear idea of their research interests and goals, with significant theoretical or practical knowledge of a chosen field. An MSc by full-time research provides an excellent training in laboratory research and a strong grounding for further study at the level of PhD. Initial enquires should be made directly to Dr. Paul Skehel Paul.Skehel@ed.ac.uk.

MSc students will work full-time on their research project with the additional option of taking selected transferable skills courses. Each student will have two supervisors. After 3 months study a short presentation and report of completed and proposed work will be made to the supervisors, and at the end of the year a dissertation will be submitted and assessed for the award of MSc.

If you're interested in applying for the MSc by Research, please review the Research Profiles and recent publications from members of the CDBS:

A preliminary email briefly outlining your research interests and identifying up to three potential supervisors from the CDBS should be sent directly to Dr Paul Skehel. If appropriate, students will then be invited to submit a short 400-word general outline for a potential MSc project with specific supervisors. Applicants progressing to this point will then be guided to make a formal application on this page. Please note that the MSc by Research has additional programme costs of £5,000.

Students have access to state-of-the-art laboratories and equipment for all aspects of modern molecular, cellular and systems biology to facilitate their research objectives. These are located both within the Central area campus and at the Little France campus. Extensive collaborations exist with the wider biomedical and clinical communities helping ensure you are supported in a world-class research environment.

Research is mainly funded by grants from the MRC, the BBSRC and industrial companies.

These entry requirements are for the 2024/25 academic year and requirements for future academic years may differ. Entry requirements for the 2025/26 academic year will be published on 1 Oct 2024.

A UK 2:1 honours degree or its international equivalent.

International qualifications

Check whether your international qualifications meet our general entry requirements:

English language requirements

Regardless of your nationality or country of residence, you must demonstrate a level of English language competency at a level that will enable you to succeed in your studies.

English language tests

We accept the following English language qualifications at the grades specified:

  • IELTS Academic: total 6.5 with at least 6.0 in each component. We do not accept IELTS One Skill Retake to meet our English language requirements.
  • TOEFL-iBT (including Home Edition): total 92 with at least 20 in each component. We do not accept TOEFL MyBest Score to meet our English language requirements.
  • C1 Advanced (CAE) / C2 Proficiency (CPE): total 176 with at least 169 in each component.
  • Trinity ISE: ISE II with distinctions in all four components.
  • PTE Academic: total 62 with at least 59 in each component.

Your English language qualification must be no more than three and a half years old from the start date of the programme you are applying to study, unless you are using IELTS, TOEFL, Trinity ISE or PTE, in which case it must be no more than two years old.

Degrees taught and assessed in English

We also accept an undergraduate or postgraduate degree that has been taught and assessed in English in a majority English speaking country, as defined by UK Visas and Immigration:

We also accept a degree that has been taught and assessed in English from a university on our list of approved universities in non-majority English speaking countries (non-MESC).

If you are not a national of a majority English speaking country, then your degree must be no more than five years old* at the beginning of your programme of study. (*Revised 05 March 2024 to extend degree validity to five years.)

Find out more about our language requirements:

Additional programme costs

Please note that the MSc by Research has additional programme costs of £5,000.

Living costs

You will be responsible for covering living costs for the duration of your studies.

Tuition fees

AwardTitleDurationStudy mode
PhDIntegrative Physiology (Biomedical Sciences - Centre for Discovery Brain Sciences)3 YearsFull-timeTuition fees
PhDIntegrative Physiology (Biomedical Sciences - Centre for Discovery Brain Sciences)6 YearsPart-timeTuition fees
MScRIntegrative Physiology (Biomedical Sciences - Centre for Discovery Brain Sciences)1 YearFull-timeTuition fees
MScRIntegrative Physiology (Biomedical Sciences - Centre for Discovery Brain Sciences)2 YearsPart-timeTuition fees

Featured funding

UK government postgraduate loans

If you live in the UK, you may be able to apply for a postgraduate loan from one of the UK's governments.

The type and amount of financial support you are eligible for will depend on:

  • your programme
  • the duration of your studies
  • your residency status.

Programmes studied on a part-time intermittent basis are not eligible.

Other funding opportunities

Search for scholarships and funding opportunities:

  • Edinburgh Medical School: Biomedical Sciences
  • University of Edinburgh
  • 1 George Square
  • Central Campus
  • Edinburgh
  • EH8 9JZ

We encourage you to apply at least one month prior to entry so that we have enough time to process your application. If you are also applying for funding or will require a visa then we strongly recommend you apply as early as possible.

You must submit two references with your application.

Before making your application, you must make contact with a potential supervisor to discuss your research proposal. Further information on making a research degree application can be found on the College website:

You will be formally interviewed (in person, by video-conferencing or Skype).

Find out more about the general application process for postgraduate programmes:

Further information

  • Edinburgh Medical School: Biomedical Sciences
  • University of Edinburgh
  • 1 George Square
  • Central Campus
  • Edinburgh
  • EH8 9JZ