1974 D.Phil. University of Sussex
1975 MRC Researc Scientist, MIMR London
1977 Lecturer University of Edinburg
1995 Senior Lecturer University of Edinburgh
Lectures in Evolution in Action 2 and Genes & Gene Action 2
Course Organiser of Developmental Biology 3
Zoology Honours Programme Organiser
I am a Developmental Biologist, mainly interested in spatial pattern formation in insects and in evolution of the mechanisms involved. Past research projects have included insect limb regeneration; segmentation in insect embryos; and the control of growth and size in relation to temperature. My current research concerns the specification of colour patterns on developing butterfly and moth wings.