1981 PhD in Immunology, Department of Pathology, University of Cambridge
1981-1984 Postdoctoral Research Assistant, Department of Pathology, University of Cambridge
1984-1987 Postdoctoral Research Fellow, Dept. of Medicine, Stanford University, U.S.A.
1987-1991 Research Scientist, ICRF, Tumour Immunology Unit, University College London
1991-2007 Senior Research Scientist, MRC National Institute for Medical Research, Mill Hill, London
2007-present Professor of Immune Cell Biology, University of Edinburgh